The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
Young amphibians generally undergo metamorphosis from an aquatic larval form with gills to an air-breathing adult form with lungs. Amphibians use their skin as a secondary respiratory interface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require access to water bodies to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators to habitat conditions; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from tetrapodomorph sarcopterygians (lobe-finned fish with articulated limb-like fins) that evolved primitive lungs, which were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (predecessors of mammals). The origin of modern lissamphibians, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondys. A fourth group of lissamphibians, the Albanerpetontidae, became extinct around 2 million years ago. (Full article...)
Selected frog article
The Alytidae are a family of primitive frogs. Their common name is painted frogs or midwife toads. Most are endemic to Europe, but three species occur in northwest Africa, and a species formerly thought to be extinct is found in Israel.
This family is also known as Discoglossidae, but the older name Alytidae has priority and is now recognized by major reference works. Some researchers, though, suggest that Alytes and Discoglossus are different enough to be treated as belonging to separate families, implying resurrection of the Discoglossidae. The term "discoglossid" has also been used to refer to many primitive fossil frogs that share plesiomorphic (ancestral) similities to alytids, but that are probably not closely related. (Full article...)Selected salamander article
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Sexual selection in amphibians involves sexual selection processes in amphibians, including frogs, salamanders and newts. Prolonged breeders, the majority of frog species, have breeding seasons at regular intervals where male-male competition occurs with males arriving at the waters edge first in large number and producing a wide range of vocalizations, with variations in depth of calls the speed of calls and other complex behaviours to attract mates. The fittest males will have the deepest croaks and the best territories, with females making their mate choices at least partly based on the males depth of croaking. This has led to sexual dimorphism, with females being larger than males in 90% of species, males in 10% and males fighting for groups of females.
There is a direct competition between males to win the attention of the females in salamanders and newts, with elaborate courtship displays to keep the females attention long enough to get her interested in choosing him to mate with. Some species store sperm through long breeding seasons, as the extra time may allow for interactions with rival sperm. (Full article...)List of selected salamander articles
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Did you know? –
- ...that the Cheat Mountain salamander of West Virginia is the only vertebrate known to be unique to that state?
- ... that the Permian amphibian Trimerorhachis either brooded young in its mouth or ate them?
- ... that the Cumberland Dusky Salamander is believed to hybridize with the Ocoee Salamander?
- ... that female European cave salamanders do not normally eat while they guard their eggs for six months or more?
- ... that in hot weather, the eastern spadefoot toad (pictured) may aestivate in the deep burrow it has dug?
Selected amphibian type
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela from the group Caudata. Urodela is a scientific Latin term based on the Ancient Greek οὐρά δήλη: ourà dēlē "conspicuous tail". Caudata is the Latin for "tailed ones", from cauda : "tail".
Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders never have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults.
This group of amphibians is capable of regenerating lost limbs as well as other damaged parts of their bodies. Researchers hope to reverse engineer the regenerative processes for potential human medical applications, such as brain and spinal cord injury treatment or preventing harmful scarring during heart surgery recovery. The remarkable ability of salamanders to regenerate is not just limited to limbs but extends to vital organs such as the heart, jaw, and parts of the spinal cord, showing their uniqueness compared to different types of vertebrates. This ability is most remarkable for occurring without any type of scarring. This has made salamanders an invaluable model organism in scientific research aimed at understanding and achieving regenerative processes for medical advancements in human and animal biology. (Full article...)Selected images
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Photo: Richard BartzThe common frog (Rana temporaria) is found throughout much of Europe. Adults have a body length of 6 to 9 cm (2.4 to 3.5 in) and vary in colour, with the ability to lighten and darken their skin to match their surroundings. They will feed on any invertebrate of a suitable size and, apart from the breeding season, live solitary lives. -
Photo: Benny TrappAn Italian tree frog (Hyla intermedia) with an inflated vocal sac. Vocal sacs, flexible membranes of skin possessed by most male frogs, are generally used to amplify mating or advertisement calls. However, some species in the family Rhinodermatidae may also use the sac to store or transport their tadpoles. -
Photograph credit: RushenbRaorchestes parvulus is a small bush frog native to tropical southeastern Asia. Although the reproductive strategy of this species has not been studied, members of the family typically make a foam nest while mating in vegetation overhanging a stream. They create the foam by beating their legs, and the eggs are deposited into the nest and covered with seminal fluid before the foam hardens into a protective casing. When the eggs hatch, the young push their way out of the nest and fall into the water below.
This picture shows a male R. parvulus frog, photographed in Phu Kradueng National Park, Thailand, calling by inflating its vocal sac. -
Photograph credit: Charles James SharpThe marsh frog (Pelophylax ridibundus) is a species of amphibian native to Europe and parts of Asia. It is the largest type of frog in most of its range, growing to a snout-to-vent length of around 100 mm (4 in); tadpoles can reach up to 190 mm (7.5 in) in length, but this usually occurs in places with long winters where the tadpole has time to grow. Marsh frogs hibernate during the winter, either underwater or in burrows, and are able to use the Earth's magnetic field to locate breeding ponds. This marsh frog was photographed in Kampinos National Park, Poland. -
Photo credit: LiquidGhoulThe New England Tree Frog (Litoria subglandulosa) is a species of frog native to the streams of the New England Tablelands, New South Wales, Australia. -
Photo: Marie-Lan NguyenThe marsh frog (Pelophylax ridibundus) is a species of true frog and the largest frog native to Europe; females of this sexually dimorphic species may be up to 17 centimetres (6.7 in) long. The marsh frog feeds mainly on insects, but it also eats smaller amphibians, fish, and rodents. -
Photograph credit: Charles J. SharpMantella baroni, commonly known as Baron's mantella, variegated golden frog or Madagascar poison frog, is a species of frog in the family Mantellidae. It is native to Madagascar and is seen here in Ranomafana National Park in the southeastern part of the country. The species was described in 1888 by George Albert Boulenger, who named it after its collector, Richard Baron. It has been classified as a least-concern species by the International Union for Conservation of Nature due to its relatively wide distribution, but is threatened by habitat loss. The frog's diet includes mites, which cause it to accumulate a high alkaloid concentration in its skin, making it toxic. The bright colours may serve as a warning sign to potential predators of the frog's toxicity. -
Photograph: Brian GratwickeRabbs' fringe-limbed treefrog (Ecnomiohyla rabborum) is a large species of frog originally found in the forest canopies of central Panama. Only discovered in 2005, the species is thought to be extinct in the wild; only one specimen, a male at the Atlanta Botanical Garden, survives. -
Photo credit: FroggydarbA specimen from the "hybrid zone" of the Leaf Green Tree Frog (Litoria phyllochroa) and the Southern Leaf Green Tree Frog (L. nudidigitus), showing physical characteristics of both species. These small stream-dwelling frogs (averaging only 40mm in length), are native to eastern Australia and occur together south of Sydney. The two species are differentiated only by distribution, call and slight differences in flank markings. -
Photograph: Brian GratwickeThe Panamanian golden frog (Atelopus zeteki) is a rare species of toad endemic to Panama. First described by Emmett Reid Dunn in 1933, this species was found close to mountain streams on the eastern side of the Tabasará mountain range in Coclé and Panamá Provinces, but is now critically endangered and possibly extinct in the wild. -
Photo credit: Carey James BalboaThe Red-eyed Tree Frog (Agalychnis callidryas) is a small-sized tree frog, reaching lengths of about 5 to 7 centimeters (3 inches), native to Neotropical rainforests in Central America. They are not poisonous and rely on camouflage to protect them. During the day, they remain motionless with their colorful parts hidden. Thus, they appear almost completely green, and well hidden among the foliage. -
Photograph: Benny TrappThe Spanish painted frog (Discoglossus jeanneae) is a species of frog in the family Alytidae. Endemic to Spain, it mostly lives in open areas, pine groves and shrublands. It feeds mostly on insects and worms. -
In the life cycle of a frog, a female lays her eggs in a shallow pond or creek, where they will be sheltered from the current and from predators. The eggs, known as frogspawn hatch into tadpoles. The tadpoles develop gradually into adolescent froglets and finally the froglet develops into an adult frog. -
A male Dendropsophus microcephalus frog, displaying its vocal sac—membranes of skin under the throat or on the corner of the mouth that distend to amplify its call. The species lives in moist areas in most of Central America and northern South America. -
The fire salamander (Salamandra salamandra) is a European salamander species with a long lifespan. These nocturnal animals generally eat various insects, spiders, earthworms and slugs, but they also occasionally eat newts and young frogs.
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Selected toad article
Selected caecilian article
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